Johnson, R. A. 1981. Application of the guild concept to enviromental impact analysis of terrestrial vegetation. Journal of Environmental Management 13:205-222.

Johnson, R. A., M. F. Willson, J. N. Thompson, and R. I. Bertin. 1985. Nutritional values of wild fruits and consumption by migrant frugivorous birds. Ecology 66:819-827.

Johnson, R. A. 1986. Intraspecific resource partitioning in the bumblee bees Bombus ternarius and B. pennsylvanicus. Ecology 67:133-138.

Rissing, S. W., R. A. Johnson, and G. B. Pollock. 1986. Natal nest distribution and pleometrosis in the leafcutter ant Acromyrmex versicolor. Psyche 93:177-186.

Johnson, R. A., S. W. Carothers, and T. J. McGill. 1987. Demography of feral burros in the Mohave Desert. Journal of Wildlife Management 51:616-620.

Johnson, R.A., and S. W. Rissing. 1990. Comparative learning and memory of two sympatric seed-harvester ants. Pp. 552-553 In: Social insects and the environment, G.K. Veeresh, B. Mallik, and C.A. Vikaktamath (eds.).   Oxford and IBH Publishing Co., PVT, LTD., New Delhi, India, 765 pp.

Johnson, R. A. 1992. Pollination and reproductive ecology of acuña cactus, Echinomastus erectrocentrus var. acunensis. International Journal of Plant Sciences 153:400-408.

Johnson, R. A. 1991. Comparative foraging efficiency of two species of desert seed-harvester ants: effects of learning and memory. Ecology 72:1408-1419.

Johnson, R. A. 1992. Soil texture as an influence on the distribution of the desert seed-harvester ants Pogonomyrmex rugosus and Messor pergandei. Oecologia 89:118-124.

Johnson, R. A., and S. W. Rissing. 1993. Breeding biology of the leaf-cutter ant Acromyrmex versicolor (Pergande) (Hymenoptera: Formicidae). Journal of the Kansas Entomological Society 66:127-128.

Johnson, R. A., M. A. Baker, D. J. Pinkava, and G. A. Ruffner. 1993. Seedling establishment, mortality, growth, and flower production in acuña cactus, Echinomastus erectrocentrus var. acunensis. Pp. 170-180, In: Sivinski, R., and K. Lightfoot (eds.). Proceedings of the Southwestern Rare and Endangered Plant Conference, Santa Fe, New Mexico. 

Lighton, J. R. B., D. A. Garrigan, F. D. Duncan, and R. A. Johnson. 1993. Spiracular control of respiratory water loss in female alates of the harvester ant Pogonomyrmex rugosus. Journal of Experimental Biology 179:233-244.

Johnson, R. A., S. W. Rissing, and P. R. Killeen. 1994. Differential learning and memory by two co-occurring ant species. Insectes Sociaux 41:165-177.

Johnson, R. A. 1994. Distribution and natural history of the workerless inquiline ant Pogonomyrmex anergismus Cole (Hymenoptera: Formicidae). Psyche 101:257-262.

Johnson, R. A., J. D. Parker, and S. W. Rissing. 1996. Rediscovery of the workerless inquiline ant Pogonomyrmex colei and additional notes on natural history (Hymenoptera: Formicidae). Insectes Sociaux 43:69-76.

Johnson, R. A. 1998. Foundress survival and brood production in the desert seed-harvester ants Pogonomyrmex rugosus and P. barbatus. Insectes Sociaux 45:255-266.

Johnson, R. A. 1998. Desiccation resistance in North American seed- harvester ants: local and latitudinal patterns in body size and habitat. Pg. 521, In: Schwarz, M. P. and Hogendoorn, K. (eds.). Social insects at the turn of the millenium. Proceedings of the XIII International Congress    of the IUSSI.

Rissing, S. W., R. A. Johnson, and J. W. Martin. 2000. Colony founding behavior of some desert ants: geographic variation in metrosis. Psyche 103:95-101.

Johnson, R. A. 2000. Water loss in desert ants: caste variation and the effect of cuticle abrasion. Physiological Entomology 25:48-53.

Johnson, R. A. 2000. Habitat segregation based on soil texture and body size in the seed-harvester ants Pogonomyrmex barbatus and P. rugosus. Ecological Entomology 25:403-412.

Johnson, R. A. 2000. Reproductive biology of the seed-harvester antsMessor julianus (Pergande) and Messor pergandei (Mayr) (Hymenoptera: Formicidae) in Baja California, Mexico. Journal of Hymenoptera Research 9:377-384.

Johnson, R.A. 2000. Seed-harvester ants (Hymenoptera: Formicidae) of North America: an overview of ecology and biogeography. Sociobiology 35:89-122.

Baker, M.A. and R.A. Johnson. 2000. Morphometric analysis of Escobaria sneedii var. sneedii, E. sneedii var. leei and E. guadalupensis (Cactaceae). Systematic Botany 25:577-587.

Johnson, R. A. 2001. Biogeography and community structure of North American seed-harvester ants. Annual Review of Entomology 46:1-29.

Johnson, R.A. 2002. Semi-claustral colony founding in the seed-harvester ant Pogonomyrmex californicus: a comparative analysis of colony founding strategies. Oecologia 132:60-67.

Cahan, S.H., J.D. Parker, S.W. Rissing, R.A. Johnson, T.S. Polony, M.D. Weiser, and D.R. Smith. 2002. Extreme genetic differences between queens and workers in hybridizing Pogonomyrmex harvester ants. Proceedings of the Royal Society of London, B 269:1871-1877.

Julian, G.E., J.H. Fewell, J. Gadau, R.A. Johnson, and D. Larrabee. 2002. Genetic determination of the queen caste in an ant hybrid zone. Proceedings of the National Academy of Sciences USA 99:8157-8160.

Johnson, R.A. and P.S. Ward. 2002. Biogeography and endemism of ants in Baja California: a first overview. Journal of Biogeography 29:1009-1026.

Hahn, D.A., R.A. Johnson, N.A. Buck, and D.E. Wheeler. 2004. Storage protein content as a functional marker for colony founding strategies: a comparative study within the harvester ant genus, Pogonomyrmex. Physiological and Biochemical Zoology 77:100-108.

Johnson, R.A. 2004. Colony founding by pleometrosis in the semi-claustral seed-harvester ant Pogonomyrmex californicus (Hymenoptera: Formicidae). Animal Behaviour 68: 1189-1200.

Gibbs, A.G. and R.A. Johnson. 2004. Discontinuous ventilation by insects: the chthonic hypothesis does ot hold water. Journal of Experimental Biology 207:3477-3482.

Johnson, R.A. and A.G. Gibbs. 2004. Effects of mating status on cuticular hydrocarbons, metabolism and water balance in the desert harvester ant Pogonomyrmex barbatus. Journal of Insect Physiology 50:943-953.

Johnson, R.A. 2006. Capital and income breeding and the evolution of colony founding strategies in ants. Insectes Sociaux, 53: 316-322.

Johnson, R.A. 2006. Biogeographical parallels between plants and ants in North American deserts (Hymenoptera: Formicidae; Spermatophyta). Myrmecological News, 8: 209-218.

Anderson, K.E., J. Gadau, B.M. Mott, R.A. Johnson, A. Altamirano, C. Strehl, and J.H. Fewell. 2006. Distribution and evolutionary history of genetic caste determination in seed-harvester ants Pogonomyrmex barbatus and P. rugosus. Ecology, 87:2171-2184.

Kronauer, D.J.C., R.A. Johnson and J.J. Boomsma. 2007. The evolution of multiple mating in army ants. Evolution 61: 413-422.

Johnson, R,.A., C.T. Holbrook, C. Strehl and J. Gadau. 2007. Population and colony structure and morphometrics in the queen dimorphic harvester ant, Pogonomyrmex pima. Insectes Sociaux 54: 77-86.

Holbrook, C.T., C.P. Strehl, R.A. Johnson and J. Gadau. 2007. Low queen mating frequency in the seed-harvester ant Pogonomyrmex (Ephebomyrmex ) pima : implications for the evolution of polyandry. Behavioral Ecology and Sociobiology 62: 229-236

Rabeling, C., S.P. Cover, R.A. Johnson and U.G. Mueller. 2007. A review of the North American species of the fungus-gardening ant genus Trachymyrmex (Hymneoptera: Formicidae) . Zootaxa 1644: 1-53

Rabeling, C., S.P. Cover, R.A. Johnson and U.G. Mueller. 2007. A review of the North American species of the fungus-gardening ant genus Trachymyrmex (Hymenoptera: Formicidae). Zootaxa 1664:153.

Oettler, J. and R.A. Johnson. 2009. The old ladies of the seed harvester ant Pogonomyrmex rugosus : foraging performed by two groups of workers .Journal of Insect Behavior 22: 217-226.

Johnson, R.A. and R.P. Overson. 2009. A new North American species of Pogonomyrmex (Hymenoptera: Formicidae) from the Mohave Desert of eastern California and western Nevada . Journal of Hymenoptera Research 18: 305-314.

Johnson, R.A. 2010. Independent colony founding by ergatoid queens in the ant genus Pogonomyrmex: queen foraging provides an alternative to dependent colony founding . Insectes Sociaux 57: 169-176

Sirviö, A. P. Pamilo, R.A. Johnson, R.E. Page Jr, and J. Gadau. 2011. Origin and evolution of the dependent lineages in the genetic caste determination system of Pogonomyrmex ants. Evolution 65: 869-884.

Johnson, R.A., A. Kaiser, M. Quinlan, and W. Sharp. 2011. Effect of cuticular abrasion and recovery on water loss rates in queens of the desert harvester ant Messor pergandei. Journal of Experimental Biology, 214: 3495-3506.

Peeters C., R.A. Keller, and R.A. Johnson. 2012. Selection against aerial dispersal in ants: two non-flying queen phenotypes in Pogonomyrmex laticeps. PLoS ONE 7(10): e47727. doi:10.1371/journal.pone.0047727.

Plowes, N.J.R., R. A. Johnson, and B. Hölldobler. 2012. Foraging behavior in the ant genus Messor (Hymenoptera: Formicidae: Myrmicinae). Myrmecological News, 18:33-49.

Johnson, R.A., R.P. Overson, and C. Moreau. 2013. A new species of seed-harvester ant, Pogonomyrmex hoelldobleri (Hymenoptera: Formicidae), from the Mohave and Sonoran Deserts of North America. Zootaxa,3646 (3): 201-227.

Hölldobler, B., N.J.R. Plowes, R.A. Johnson, U. Nishshanka, C. Liu, and A.B. Attygalle. 2013. Pygidial gland chemistry and potential alarm-recruitment function in column foraging, but not solitary, Nearctic Messor harvesting ants (Hymenoptera: Formicidae: Myrmicinae). Journal of Insect Physiology, 59: 863-869.

Plowes, N.J.R., T. Colella, R.A. Johnson and B. Hölldobler. Chemical communication during foraging in the harvesting ants Messor pergandei and M . andrei . Journal of Comparative Physiology A, 200: 129-137.

Favé, M.J., R.A. Johnson, S.P. Cover, S. Handschuh, B. Metscher, G.B. Müller, S. Gopalan, and E. Abouheif. 2015. Climate change drives novelties in Sky Island ant populations. Proceedings of the National Academy of Sciences, in review.

Johnson, R.A. & S.P. Cover. 2015. A taxonomic revision of the seed-harvester ant genus Pogonomyrmex (Hymenoptera: Formicidae) on the island of Hispaniola, Zootaxa, 3972 (2): 231-249.

Johnson, R.A. 2015. A taxonomic revision of South American species of the seed-harvester ant genus Pogonomyrmex (Hymenoptera: Formicidae). Part I. Zootaxa, 4029 (2): 1-142.

Johnson, R.A. and C.S. Moreau. 2016. A new ant genus from southern Argentina and southern Chile, Patagonomyrmex (Hymenoptera: Formicidae). Zootaxa, 4139 (1): 1-31.

Amorim, D.S., C.M.D. Santos, F. Krell, A. Dubois, S.S. Nihei, O.M.P. Oliveira, A. Pont, H. Song, V.K. Verdade, D.A. Fachin, B. Klassa, C.J. Lamas, S.S. Oliveira, C.J.B. de Carvalho, C.A. Mello-Patiu, E. Hajdu, M.S. Couri, V.C. Silva, R.S. Capellar, R.L. Falaschi, R. Feitosa, L. Prendini, J.P. Pombal Jr., F. Fernández, R.M. Rocha, J.E. Lattke, U. Caramaschi, M. Duarte, A.C. Marques, R.E. Reis, O. Kurina, D.M. Takiya , M. Tavares, D.S. Fernandes, .L.Franco, F. Cuezzo , D. Paulson, B. Guénard, B.C. Schlick-Steiner, W.Arthofer, F.M. Steiner , B.L. Fisher , R.A. Johnson , T.D. Delsinne , D.A. Donoso, P.R. Mulieri, L.D. Patitucci & D. Grimaldi 2016. Timeless standardsfor species delimitation: A critique of the use of images as types. Zootaxa, 4137 (1):121-128.

Johnson, R.A. and R.P. Overson. 2017. Population and colony structure and morphometrics in the queen dimorphic little black ant, Monomorium AZ-02, with a review of queen phenotypes in the genus Monomorium. PLoS ONE 12 (7): e0180595.

Mueller, U., H. Ishak, S. Bruschi, C. Smith, J. Herman, S. Solomon, A. Mikheyev, C. Rabeling, J. Scott, M. Cooper, A. Rodrigues, A. Ortiz, C.R. Brandão, J. Lattke, F. Pagnocca, S. Rehner, T.R. Schultz, H. Vasconcelos, R.Adams, M. Bollazzi, R. Clark, A. Himler, J. LaPolla, I. Leal, R. Johnson, F.Roces, J. Sosa-Calvo, R. Wirth and M. Bacci. Biogeography of mutualistic fungi cultivated by leafcutter ants. Molecular Ecology, in review.


Johnson, R. A. 1996. Arizona ants. Arizona Wildlife Views. June, pp. 2-5.